The invert function converts every color in the selection to it's opposite - for example, black becomes white, yellow becomes blue etc. The effect this create is similar the a photograph negative. You can see what the opposite of all colors is by looking at the GTK color selector (double-click the foreground or background color, and select the "GTK" tab). The opposite color is the one at the opposite side of the color wheel.

If no selection is made, the entire layer is inverted.

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